Because Exchange auto-complete cache uses X.500 addresses for e-mail sent to addresses within the Exchange organization, it will bounce back messages from a re-created mailbox even after you give the user account all the old SMTP addresses. This is because the old X.500 address in the auto-complete cache is missing, and this causes Exchange to reject the messages. Ben Lye explains how to solve this common problem.
Source : Exchange E-mail Addresses and the Outlook Address Cache – Simple Talk